Know more

About cookies

What is a "cookie"?

A "cookie" is a piece of information, usually small and identified by a name, which may be sent to your browser by a website you are visiting. Your web browser will store it for a period of time, and send it back to the web server each time you log on again.

Different types of cookies are placed on the sites:

  • Cookies strictly necessary for the proper functioning of the site
  • Cookies deposited by third party sites to improve the interactivity of the site, to collect statistics

Learn more about cookies and how they work

The different types of cookies used on this site

Cookies strictly necessary for the site to function

These cookies allow the main services of the site to function optimally. You can technically block them using your browser settings but your experience on the site may be degraded.

Furthermore, you have the possibility of opposing the use of audience measurement tracers strictly necessary for the functioning and current administration of the website in the cookie management window accessible via the link located in the footer of the site.

Technical cookies

Name of the cookie


Shelf life

CAS and PHP session cookies

Login credentials, session security



Saving your cookie consent choices

12 months

Audience measurement cookies (AT Internet)

Name of the cookie


Shelf life


Trace the visitor's route in order to establish visit statistics.

13 months


Store the anonymous ID of the visitor who starts the first time he visits the site

13 months


Identify the numbers (unique identifiers of a site) seen by the visitor and store the visitor's identifiers.

13 months

About the AT Internet audience measurement tool :

AT Internet's audience measurement tool Analytics is deployed on this site in order to obtain information on visitors' navigation and to improve its use.

The French data protection authority (CNIL) has granted an exemption to AT Internet's Web Analytics cookie. This tool is thus exempt from the collection of the Internet user's consent with regard to the deposit of analytics cookies. However, you can refuse the deposit of these cookies via the cookie management panel.

Good to know:

  • The data collected are not cross-checked with other processing operations
  • The deposited cookie is only used to produce anonymous statistics
  • The cookie does not allow the user's navigation on other sites to be tracked.

Third party cookies to improve the interactivity of the site

This site relies on certain services provided by third parties which allow :

  • to offer interactive content;
  • improve usability and facilitate the sharing of content on social networks;
  • view videos and animated presentations directly on our website;
  • protect form entries from robots;
  • monitor the performance of the site.

These third parties will collect and use your browsing data for their own purposes.

How to accept or reject cookies

When you start browsing an eZpublish site, the appearance of the "cookies" banner allows you to accept or refuse all the cookies we use. This banner will be displayed as long as you have not made a choice, even if you are browsing on another page of the site.

You can change your choices at any time by clicking on the "Cookie Management" link.

You can manage these cookies in your browser. Here are the procedures to follow: Firefox; Chrome; Explorer; Safari; Opera

For more information about the cookies we use, you can contact INRAE's Data Protection Officer by email at or by post at :


24, chemin de Borde Rouge -Auzeville - CS52627 31326 Castanet Tolosan cedex - France

Last update: May 2021

Menu Logo Principal logo Université Clermont Auvergne & associés

Human Nutrition Unit

Zone de texte éditable et éditée et rééditée

Team members



Fafournoux Pierre

Senior scientist CNRS

Team leader

Regulation of physiological functions by amino acid availability ; protein and amino acids homeostasis; eIF2a dependent signaling

Phone +33 (0)4 73 62 45 62   

E-mail pierre.fafournoux[at]


MD-PhD, Clinic Leader, Hospital Practitioner, Clermont-Fd Hospital

Nephrology, Kidney disease, Extrarenal Purification Muscle Atrophy, Proteolytic systems, Biomarkers



Averous Julien


Amino acids, polyphenols, translation, GCN2, ATF4, mTORC1, Integrated Stress Response

Phone +33 (0)4 73 75 23 61

Phone +33 (0)4 73 62 42 02    

E-mail julien.averous[at]


Post-Doctoral Fellow

Molecular and cellular biology, GCN2/eIF2a/ATF4, Gene therapy, Viral vectors, Transgenic mice, Gene regulation by nutrition


Bruhat Alain

Senior scientist

Amino acids, GCN2, ATF4, Integrated Stress Response, Autophagy, muscle atrophy, Bioluminescent mouse, Gene therapy


Phone +33 (0)4 73 62 40 00

E-mail fida.azar[at]

Phone + 33 (0)4 73 62 41 50   

E-mail alain.bruhat[at]   


Carraro Valérie

Engineer assistant

Molecular and cell biology, Transgenic mice, Animal experimentation, Health and safety



Claustre Agnes

Engineer assistant

Molecular and Cell Biology, Electroporation, Animal Experimentation, Human, Ubiquitin-Proteasome, UBE2, E3-Ligase, Muscle atrophy


Phone + 33 (0)4 73 62 40 44     

E-mail valerie.carraro[at]

Phone +33 (0)4 73 62 48 45     

E-mail agnes.claustre[at]


Combaret Lydie

Senior scientist

Muscle atrophy, Physical inactivity, Mitochondria, Autophagy, Ubiquitin-Proteasome, atrophy-induced models


Coudy Cécile

Engineer assistant

Histology, Imaging, Aging, Muscle atrophy, Pathology, Human, Animal models, Health and safety

Phone +33 (0)4 73 62 48 24   

E-mail lydie.combaret[at]   

Phone /Office +33 (0)4 73 62 42 07 /Histo lab +33 (0)4 73 62 48 68

E-mail cecile.coudy-gandilhon[at]


Coralie Delabrise

Ph. Student

Muscle atrophy, Biomarkers, Mechanisms, Human, Cellular models 

FAURE Audrey

Faure Audrey


Cell and molecular biology, Gene regulation by nutrition, Viral vectors

E-mail coralie.delabrise[at]

Phone +33 (0)4 73 62 44 98     

E-mail audrey.faure[at] 


Ph. Student

Brown bear hibernation, Radioresistance, Physiology, Cellular models, Molecular Biology

HENG Anne-Elisabeth

Heng Anne-Elisabeth

Pr. Nephrology, CHU Clermont-Fd

Nephrology, Kidney failure, Muscle atrophy, Proteolytic system, Biomarkers

E-mail guillaume.fourneaux[at]

Phone +33 (0)4 73 75 14 25


Céline Jousse

Scientist CNRS

Nutritional programming, Epigenetic, Methylation, Metabolic disease, Markers

LEFAI Etienne

Lefai Etienne

Senior scientist

Muscle atrophy, Bear hibernation, Physiology, Radioprotection

Phone +33 (0)4 73 62 49 39   

E-mail celine.jousse[at]    

Phone +33 (0)4 73 62 47 44   

E-mail etienne.lefai[at]  

MAURIN Anne-Catherine

Maurin Anne-Catherine


Amino acid, Stress signaling, GCN2, eIF2a, ATF4, Food intake, Anorexia, Autophagy

PAPET Isabelle

Papet Isabelle


Sulfur amino acids, Amino acid availability, Muscle wasting, Nutritional strategies, Medication and amino acid/protein homeostasis, Inter-organ relationships


Phone +33 (0)4 73 62 41 43   

E-mail anne-catherine.maurin[at]

Phone +33 (0)4 73 62 42 01     

E-mail isabelle.papet[at]    

PARRY Laurent

Parry Laurent

Engineer assistant

Molecular and cellular biology, Animal expérimentation, Integrated Stress Response, Nutritional programming, Methylation


POLGE Cécile

Polge Cécile


Ubiquitin-proteasome system (UPS), E3 ligase, MuRF1, TRIM63, UBE2, Muscle atrophy


Phone+33 (0)4 73 62 47 69   

E-mail laurent.parry[at]

Phone + 33 (0)4 73 62 42 18   

E-mail cecile.polge[at]


Maxime Raucroy

PhD. Student

GCN2/ATF4 signalling, Essential amino acid restriction, Plant proteins, Nutrition



PhD. Student

TGFb/BMP signaling, Bear hibernation, Cellular models, Muscle atrophy, Cell and molecular biology

E-mail maxime-ludovic-pierre.raucroy[at]       

  E-mail chloe.richard2[at]      


Taillandier Daniel

Senior scientist

Ubiquitin-proteasome system (UPS), Ubiquitin Conjugating Enzymes, E3 ligases, MuRF1/TRIM63, Muscle atrophy, Biomarkers

VITURET Cyrielle

Vituret Cyrielle


Viral vectors, Molecular and cell biology, Gene regulation by nutrition (Nutrireg), translation to gene therapy

Phone +33 (0)4 73 62 48 44   

E-mail daniel.taillandier[at] 

Phone +33 (0)4 73 62 42 78   

E-mail cyrielle.vituret[at]   


  • Mélodie Malige, Technician FRM project (2018-2021)
  • Jérémy Tournayre, Identification of epigenetic markers of predisposition to obesity induced by perinatal nutritional stress (Thèse 2019). Now Engineer at INRAE.
  • Florent Mesclon, Resistance of cancer cells to amino acid deficiency (Ph.D. 2017). Now Engineer in Intelligency Business.
  • Marine Gueugneau, Human muscle alterations associated with aging and metabolic syndrome (Ph.D. 2014). Now Scientist INRAE (Clermont-Ferrand, France)
  • Laëtitia Théron, Molecular imaging of sarcopenia  (Post-Doctoral fellow 2014). Now Scientist INRAE (Clermont-Ferrand, France)
  • Yuki Muranishi, Impact of perinatal protein undernutrition on fertility in male offspring (Post Doc 2014). Assistant professor, Obihiro University of Agriculture and Veterinary Medecin (Japon)
  • Rosa Leulmi, Role of the enzyme UBE2B in the development of muscle atrophy (Thèse 2013)
  • Lamia Slimani, Mechanisms of muscle wasting during physical inactivity (Ph.D 2012). Now Professeur Assistant, Université Mohammed VI des Sciences de la Santé (Casablanca, Maroc).
  • Huijuan Wang, ge-related structural and compositional variations of mouse skeletal muscle (Ph.D. 2011). Now Engineer in Intelligency Business, Singapore
  • Justine Letienne Interaction between the 19S regulatory complex and E3 ligase Mdm2: role and mechanisms (Thèse 2010)
  • Cedric Chaveroux, Chargé de Recherche CNRS (Thèse 2009) (Centre de lutte contre le cancer Léon Berard, Lyon, France)
  • Emilie Vazeille, Mechanisms of muscle wasting during physical inactivity (Ph.D. 2008). Now Engineer, CHU Clermont-Ferrand, France
  • Yoan Chérasse, Study of the molecular mechanisms involved in the regulation of CHOP gene transpcription during amino acid deficiency (Ph.D. 2007). Now Scientist, Sakurai Laboratory, University of Tsukuba, Japan
  • Sophie Ventadour, Purification of proteasome substrates by affinity chromatography (Thèse 2006). Now Schoolteatcher
  • Isabelle Piec,  Proteomics of muscle aging in lou/c/jall rat (Ph.D. 2005). Now Senior Research Associate, Norwich Medical School, Faculty of Medicine and Health Sciences, University of East Anglia, UK
  • Amina Tassa, Regulation of muscle proteolysis by amino acids (Ph.D. 2003). Now Professor, Collin County College, Texas