Know more

About cookies

What is a "cookie"?

A "cookie" is a piece of information, usually small and identified by a name, which may be sent to your browser by a website you are visiting. Your web browser will store it for a period of time, and send it back to the web server each time you log on again.

Different types of cookies are placed on the sites:

  • Cookies strictly necessary for the proper functioning of the site
  • Cookies deposited by third party sites to improve the interactivity of the site, to collect statistics

Learn more about cookies and how they work

The different types of cookies used on this site

Cookies strictly necessary for the site to function

These cookies allow the main services of the site to function optimally. You can technically block them using your browser settings but your experience on the site may be degraded.

Furthermore, you have the possibility of opposing the use of audience measurement tracers strictly necessary for the functioning and current administration of the website in the cookie management window accessible via the link located in the footer of the site.

Technical cookies

Name of the cookie


Shelf life

CAS and PHP session cookies

Login credentials, session security



Saving your cookie consent choices

12 months

Audience measurement cookies (AT Internet)

Name of the cookie


Shelf life


Trace the visitor's route in order to establish visit statistics.

13 months


Store the anonymous ID of the visitor who starts the first time he visits the site

13 months


Identify the numbers (unique identifiers of a site) seen by the visitor and store the visitor's identifiers.

13 months

About the AT Internet audience measurement tool :

AT Internet's audience measurement tool Analytics is deployed on this site in order to obtain information on visitors' navigation and to improve its use.

The French data protection authority (CNIL) has granted an exemption to AT Internet's Web Analytics cookie. This tool is thus exempt from the collection of the Internet user's consent with regard to the deposit of analytics cookies. However, you can refuse the deposit of these cookies via the cookie management panel.

Good to know:

  • The data collected are not cross-checked with other processing operations
  • The deposited cookie is only used to produce anonymous statistics
  • The cookie does not allow the user's navigation on other sites to be tracked.

Third party cookies to improve the interactivity of the site

This site relies on certain services provided by third parties which allow :

  • to offer interactive content;
  • improve usability and facilitate the sharing of content on social networks;
  • view videos and animated presentations directly on our website;
  • protect form entries from robots;
  • monitor the performance of the site.

These third parties will collect and use your browsing data for their own purposes.

How to accept or reject cookies

When you start browsing an eZpublish site, the appearance of the "cookies" banner allows you to accept or refuse all the cookies we use. This banner will be displayed as long as you have not made a choice, even if you are browsing on another page of the site.

You can change your choices at any time by clicking on the "Cookie Management" link.

You can manage these cookies in your browser. Here are the procedures to follow: Firefox; Chrome; Explorer; Safari; Opera

For more information about the cookies we use, you can contact INRAE's Data Protection Officer by email at or by post at :


24, chemin de Borde Rouge -Auzeville - CS52627 31326 Castanet Tolosan cedex - France

Last update: May 2021

Menu Logo Principal Logo UCA

Home page


PIAF Thesis 2020 :

  • Modeling the effects of tree canopy structure on the availability of water, nutrients and microclimate


Tree canopy structure has a clear influence on resource availability (light, water, nitrogen) in natural or cultivated tree ecosystems. This structure induces emergent meteorological and microclimatic properties that can influence functional interactions between different plant strata (Gaudio et al., 2017). To optimize and predict the evolution of different functions of these tree ecosystems (microclimate, production, resource use efficiency) it is necessary to take into account these multi-stratum interactions. The hypothesis tested would be that there is (are) a tree structure(s) (neither too open nor too closed, tiered) that allow to optimize these different functions under different assumptions of climate forcing. The idea is, based on one or two case studies in the field, to parameterize and validate a model in order to test in silico the effect of different tree structures with increasing levels of complexity. The thesis will rely on the modelling platform Capsis, already used by the team and in which candidate multistratum models already exist but require development.

Gaudio N, Gendre X, Saudreau M, Seigner V, Balandier P. 2017. Agricultural and Forest Meteorology, 237-238: 71-79.


  • Study of the dynamics of the transcriptional and post-transcriptional response of Black Poplar to water deficiency

Contacts :,,

Future climate change modelling predicts global warming in Western Europe associated with an increase in the frequency and severity of extreme weather events. In this context, exploring the response of plants to water scarcity is of great importance in terms of maintaining production and even the survival of individuals in their natural range.


Black poplar is a model for the study of woody plants by the scientific community (rapid growth, sequenced and annotated genome, etc.). An initial study carried out at UMR PIAF on different genotypes of contrasting black poplars (tolerant or avoidant strategy) has made it possible to identify and analyse the role of physiological plasticity in relation to their strategy for responding to drought. The aim of this thesis is to highlight the network of transcribed genes involved in the black poplar response to drought. For this purpose, transcriptomic analysis of RNA and microRNA in kinetics will be carried out on black poplar leaves of contrasting strategies subjected to progressive hydric deficiency. Particular attention will be paid to the regulatory network of genes encoding aquaporins and to the tissue localization of candidate gene expression. This study will ultimately make it possible to decipher the key molecular mechanisms of trees' response to drought.


Garavillon-Tournayre et al. 2018. Integrated drought responses of black poplar: How important is phenotypic plasticity? Physiol. Plant. 163(1):30-44

  • Deciphering the processes of dehydration-induced cell mortality and their relationship to hydraulic failure.

Contacts :,

Xylem hydraulic failure has been identified as a ubiquitous factor in triggering drought-induced tree mortality. It results in the blockage of water transport from roots to leaves, leading to accelerated desiccation of downstream tissues. Yet, the mechanistic link between hydraulic failure and drought-induced cellular damage is still unknown. This PhD proposal aims to elucidate this link to improve our predictions of drought-induced tree mortality, focusing on three objectives:

- Analyze the relationship between cell mortality and the plant's ability to recover from drought.

- To test the relationship between cell mortality and hydraulic failure in species with different drought resistance strategies.

- Assess the effect of drought dynamics on the different processes leading to cell death.

The PhD student will be trained in new and varied techniques to assess variation in plant hydraulic functioning, cell damage and recovery capacity in different plant species exposed to different drought conditions.


- Mantova M, Herbette S, Cochard H & Torres-Ruiz JM. 2022. Hydraulic failure and tree mortality: From correlation to causation. Trends in Plant Science.

- Lemaire C, Blackman CJ, Cochard H, Menezes-Silva PE, Torres-Ruiz JM & Herbette S. 2021. Acclimation of hydraulic and morphological traits to water deficit delays hydraulic failure during simulated drought in poplar. Tree Physiology 41, 2008-2021.

  •  Root cap response to soil impedance change

Contact :

Plant roots are continuously submitted to mechanical constraints due to changing edaphic conditions (compaction, water deficit). Although the entire root is in contact with the soil, many studies have highlighted the importance of the root cap, located at the root tip, in the perception and integration of edaphic factors. Our analyses of root growth dynamics in a soil model suggest that the cap is a key zone in the perception of mechanical signals (Roué et al., 2020). The objective of this thesis is to decipher the transduction mechanisms in response to an axial force in Arabidopsis roots. We propose to take an interdisciplinary approach by combining physical and biological approaches. In a first step, we will finely analyze the deformation of root apex cells using 3D images using a light sheet microscope (SPIM for Selective Plane Illumination Microscopy), available in the laboratory. Then, we will target the key actors involved in deformation-induced transduction mechanisms. Our studies will focus on ROS and jasmonates. In plants, a number of mechanosensitive ion channels have been identified including the MSL, OSCAR, MCA and more recently PIEZZO channel families for which we will focus on phenotyping mutants.


Roué J., et al, 2020. Root cap size and shape influence responses to the physical strength of the growth medium in Arabidopsis thaliana primary roots. J. Exp. Bot., 71, 126–137.